On another note, I just spent some time looking at the animation and Black_Tiger's sprites are not representative of the full picture, the Mega Drive version has one more frame than PC-Engine with the human punch, and it has 1 more frame of animation in the Wolf's walk (I did also notice that the wolf's jump has 1 more frame on the PC-Engine, as the jump and the dash attack re-use a frame on Mega Drive)
Those sprite sheets were made by awack. I believe that they were supposed to be from the HuCard version. Separate from player frames, there's the boss growing, special effects, smoke cloud, etc that are little to no animation and very little detail in the Genesis version.
I've only checked the first stage (and that took long enough) but it seems to indicate that frame numbers have different actions winning out in either version.
The ground colour is neither suble, nor is it close to the arcade (MD's is waay closer in this area), the sprite shading is certainly more suble, but is also lacking in variety in some areas (I don't like the human's much, dragons are much better on PC-Engine though, as are the wolves IMO), the monument colour is definitely closer to the arcade version.